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Ted Evans

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Insect ecology, biological control, ecology of invasions

Professor - Biology Department

Professor

Contact Information

Office Location: BNR 245
DialPhone: 435-797-2552
SendEmail: ted.evans@usu.edu


Educational Background

PhD, Cornell University, 1980
MS, Cornell University, 1976
BA, Carleton College, 1973

Expertise

My students and colleagues and I collaborate in studies of insect ecology focused on biological control and biological invasions. Our field studies include both agricultural crops and native habitats (e.g., sagebrush communities). We study interactions among insect species (predators, parasitoids and herbivores) and host plants (including weeds and crops). We seek broadly to understand how these interactions influence population dynamics of individual species. Ultimate goals include reducing modern reliance on pesticides (insecticides and herbicides), and conserving native species in a world of increasing human population, agricultural and urban intensification, and climate change. Major topics of interest include the effects of predator diversity on prey suppression; the effects of alternative prey and other naturally occurring foods on target prey suppression; intraguild predation and other effects of invasive species on resident natural enemies and their prey; efforts to enhance predator effectiveness (conservation biological control); and effects of weather and climate change on interactions among natural enemies and their hosts. Much of the work on predators has focused on ladybird beetles (aka ladybugs; Coccinellidae), particularly as native species have been affected by spectacular successes of introduced species in colonizing and spreading throughout North America. We focus on biological control as a metapopulation process of natural enemies tracking down their pest victims across space and time. This focus highlights that success in biocontrol programs (both classical and conservation) varies widely as natural enemies are better or less able to find and contain the pest at multiple locations across often diverse landscapes. Weather and climate change further influence biocontrol success. Our current study of host-parasitoid interactions as played out in agricultural fields of northern Utah addresses these central themes.

Awards

2010 Award for Excellence in Research (Team Award), 2010

Western Association of Agricultural Experiment Station Directors

2009 APHIS Deputy Administrator's Safeguarding Award (Team Award), 2009

USDA-APHIS

2009 IPM Team Award to Western Cereal Leaf Beetle Team, 2009

Entomological Society of America, Pacific Branch

2009 Western Extension Directors' Award of Excellence (Team Award), 2009

Western Extension Directors

Outstanding Service Award, 2002

Utah Weed Control Association

Publications - Abstracts

    Publications - Books & Book Chapters

      Book Chapters

    • Hodek, I., Evans, E.W, (2012). Food Relationships: Ecology and Behaviour of the Ladybird Beetles (Coccinellidae). Wiley-Blackwell
    • Evans, E.W, Snyder, W.E, (2011). Ladybugs: Encyclopedia of Biological Invasions . University of California Press
    • Kaufman, D.W, Kaufman, G.A, Fay, P.A, Zimmerman, J.L, Evans, E.W, (1998). Animal populations and communities: Grassland dynamics: Long-term Ecological Research in Tallgrass Prairie. Oxford University Press
    • Evans, E.W, Seastedt, T.R, (1995). The relations of phytophagous invertebrates and rangeland plants. In: D.J. Bedunah and R.E. Sosebee (eds.), Wildland plants: physiological ecology and developmental morphology
    • Evans, E.W, (1994). Indirect interactions among phytophagous insects: aphids, honeydew and natural enemies. In: Leather, S.R., et al. (Eds.) Individuals, Populations, and Patterns in Ecology

    * Has not been peer reviewed

    Publications - Fact Sheets

      * Has not been peer reviewed

      Publications - Curriculum

        * Has not been peer reviewed

        Publications - Journal Articles

          Academic Journal

        • Keyser, C.A, Fernandes, E.K, Rangel, D.E, Foster, R., Jech, L.E, Reuter, K., Black, L.R, Jaronski, S., Flake, D.D, Evans, E.W, Roberts, D.W, (2017). Laboratory bioassays and field-cage trials of Metarhizium spp. isolates with field-collected Mormon crickets (Anabrus simplex). BioControl, 62, 257-268. doi: DOI 10.1007/s10526-016-9782-8
        • Evans, E.W, (2016). Biodiversity, ecosystem functioning, and classical biological control. Applied Entomology and Zoology, 51, 173-184.
        • Bolshakova, V.J, Evans, E.W, (2016). Phenology of the sagebrush defoliating moth Aroga websteri (Lepidoptera: Gelechiidae), with application to population irruptions. Annals of the Entomological Society of America, 109, 424-431.
        • Roy, H.E, Brown, P.M, Adriaens, T., Berkvens, N., Borges, I., Clusella-Trullas, S., Comont, R.F, De Clercq, P., Eschen, R., Estoup, A., Evans, E.W, Facon, B., Gardiner, M.M, Gil, A., Grez, A.A, Guillemand, T., Haelewaters, D., Herz, A., Honek, A., Howe, A.G, Hui, C., Hutchison, W.D, Kenis, M., Koch, R.L, Kulfan, J., Handley, L.L, Lombaert, E., Loomans, A., Losey, J., Lukashuk, A.O, Maes, D., Magro, A., Murray, K.M, Martin, G.S, Martinkova, Z., Minnaar, I.A, Nedved, O., Orlova-Bienkowskaua, M.J, Osawa, N., Rabitsch, W., Raven, H.P, Rondoni, G., Rorke, S.L, Ryndevich, S.K, Saethre, M., Sloggett, J.J, Soares, A.O, Stals, R., Tinsley, M.C, Vandereycken, A., van Wielink, P., Viglasova, S., Zach, P., Zakharov, I.A, Zaviezo, T., Zhao, Z., (2016). The harlequin ladybird, Harmonia axyridis: global perspectives on invasion history and ecology. Biological Invasions, 18, 997-1044.
        • Anderson, J.M, Willden, S.A, Wright, D.L, Evans, E.W, (2015). Long-term Outcomes of Population Suppression of Leafy Spurge by Insects in the Mountain Foothills of Northern Utah. American Midland Naturalist, 174, 1-13.
        • Evans, E.W, Bolshakova, V., Carlile, N.R, (2015). Parasitoid dispersal and colonization lag in disturbed habitats: biological control of cereal leaf beetle metapopulations. Journal of Applied Entomology, 139, 529-538.
        • Evans, E.W, (2015). Rapid but limited aggregation of ladybird beetles (Coleoptera: Coccinellidae) in response to sugar availability in the field. Acta Societitas Zoologicae Bohemicae, 79, 65-71.
        • Kajita, Y., Obrycki, J.J, Sloggett, J.J, Evans, E.W, Haynes, K.F, (2014). Do defensive chemicals facilitate intraguild predation and influence invasion success in ladybird beetles?. Journal of Chemical Ecology, 40, 1212-1219.
        • Evans, E.W, Carlile, N.R, Innes, M.B, Pitigala, N., (2014). Infestation of grain fields and degree-day phenology of the cereal leaf beetle (Coleoptera: Chrysomelidae) in Utah: long-term patterns. Journal of Economic Entomology/Entomological Society of America, 107, 240-249.
        • Bolshakova, V.l, Evans, E.W, (2014). Spatial and temporal dynamics of Aroga moth (Lepidoptera: Gelechiidae) populations and damage to sagebrush in shrub steppe across varying elevation. Environmental Entomology, 43, 1475-1484.
        • Evans, E.W, Carlile, N.R, Innes, M.B, Pitigala, N., (2013). Warm springs reduce parasitism of the cereal leaf beetle through phenological mismatch. Journal of Applied Entomology, 137, 383-391.
        • Hemptinne, J.L, Magro, A., Evans, E.W, Dixon, A., (2012). Body size and the rate of spread of invasive ladybird beetles in North America. Biological Invasions, 14, 595-605.
        • Losey, J.E, Perlman, J., Kopco, J., Ramsey, S., Helsler, L.S, Evans, E.W, Allee, L., Smyth, R., (2012). Potential causes and consequences of decreased body size in field populations of Coccinella novemnotata. Biological Control, 61, 98-103.
        • Evans, E.W, Comont, R.F, Rabitsch, W., (2011). Alien arthropod predators and parasitoids: interactions with the environment. BioControl, 56, 395-407.
        • Davidson, L.N, Evans, E.W, (2011). Determining diets of ladybird beetles (Coleoptera: Coccinellidae) from contents of frass.. Annals of the Entomological Society of America, 104, 675-681.
        • Evans, E.W, Soares, A.O, Yasuda, H., (2011). Invasions by ladybugs, ladybirds, and other predatory beetles. BioControl, 56, 597-611.
        • Kajita, Y., Evans, E.W, (2010). Alfalfa fields promote high reproductive rate of an invasive predatory lady beetle.. Biological Invasions, 12, 2293-2302.
        • Davidson, L.N, Evans, E.W, (2010). Frass analysis of diets of aphidophagous lady beetles (Coleoptera: Coccinellidae) in Utah alfalfa fields.. Environmental Entomology, 39, 576-582.
        • Kajita, Y., Evans, E.W, (2010). Relationships of body size, fecundity, and invasion success among predatory lady beetles (Coleoptera: Coccinellidae) inhabiting alfalfa fields. Annals of the Entomological Society of America, 103, 750-756.
        • Evans, E.W, Anderson, M.R, Bowling, P.D, (2010). Targeted sugar provision promotes parasitism of the cereal leaf beetle, Oulema melanopus. Agricultural and Forest Entomology, 12, 41-47.
        • Evans, E.W, (2009). Lady beetles as predators of insects other than Hemiptera. Biological Control, 51, 255-267.
        • Kajita, Y., Evans, E.W, (2009). Ovarian dynamics and oosorption in two species of predatory lady beetles (Coleoptera: Coccinellidae). Physiological Entomology, 34, 185-194.
        • Kajita, Y., Evans, E.W, Yasuda, H., (2009). Reproductive responses of invasive and native predatory lady beetles (Coleoptera: Coccinellidae) to varying prey availability. Environmental Entomology, 38, 1283-1292.
        • Sato, S., Yasuda, H., Evans, E.W, Dixon, A.F, (2009). Vulnerability of larvae of two species of aphidophagous ladybirds, Adalia bipunctata Linnaeus and Harmonia axyridis Pallas, to cannibalism and intraguild predation.. Entomological Science, 12, 111-115.
        • Rieder, J.P, Newbold, T.A, Sato, S., Yasuda, H., Evans, E.W, (2008). Intra-guild predation and variation in egg defence between sympatric and allopatric populations of two species of ladybird beetles.. Ecological Entomology, 33, 53-58.
        • Evans, E.W, (2008). Multitrophic interactions among plants, aphids, alternate prey and shared natural enemies - a review.. European Journal of Entomology, 105, 369-380.
        • Evans, E.W, Toler, T.R, (2007). Aggregation of polyphagous predators in response to multiple prey: ladybirds (Coleoptera: Coccinellidae) foraging in alfalfa.. Population Ecology , 49, 29-36.
        • Roberts, D.W, Rangel, D.E, Keyser, C.A, Bignayan, H.G, Dettenmaier, S.J, Fernandes, E.K, Miller, M.P, Evans, E.W, (2007). The Mormon cricket, an old threat in modern day Western USA: A search for fungal pathogens. Journal of Anhui Agricutlural University, 34:2, 141-148.
        • Snyder, W.E, Evans, E.W, (2006). Ecological effects of invasive arthropod generalist predators. Annual Review of Ecology, Evolution, and Systematics, 37, 95-122.
        • Kajita, Y., Yasuda, H., Evans, E.W, (2006). Effects of native ladybirds on oviposition of the exotic species, Adalia bipunctata (Coleoptera: Coccinellidae), in Japan. Applied Entomology and Zoology, 41, 57-62.
        • Kajita, Y., Takano, F., Yasuda, H., Evans, E.W, (2006). Interactions between introduced and native predatory ladybirds (Coleoptera, Coccinellidae): factors influencing the success of species introductions. Ecological Entomology, 31, 58-67.
        • Evans, E.W, Karren, J.B, Israelsen, C.E, (2006). Interactions over time between cereal leaf beetle (Coleoptera: Chysomelidae) and larval parasitoid Tetrastichus julis (Hymentoptera: Eulophidae) in Utah.. Journal of Economic Entomology, 99, 1967-1973.
        • Sato, S., Yasuda, H., Evans, E.W, (2005). Dropping behaviour of larvae of aphidophagous ladybirds and its effects on incidence of intraguild predation.. Ecological Entomology, 29, 27-34.
        • Toler, T.R, Evans, E.W, Tepedino, V.J, (2005). Pan-trapping for bees (Hymenoptera: Apiformes) in Utah's West Desert: the importance of color diversity.. Pan-Pacific Entomologist, 81, 103-113.
        • Evans, E.W, Gunther, D.I, (2005). The link between food and reproduction in aphidophagous predators: a case study with Harmonia axyridis (Coleoptera: Coccinellidae). European Journal of Entomology, 102, 423-430.
        • Yasuda, H., Evans, E.W, Kajita, Y., Urakawa, K., Takizawa, T., (2004). Asymmetric larval interactions between introduced and indigenous ladybirds in North America.. Oecologia, 141, 722-731.
        • Evans, E.W, (2004). Habitat displacement of native North American ladybirds by an introduced species.. Ecology, 85, 637-647.
        • Jacob, H.S, Evans, E.W, (2004). Influence of different sugars on the longevity of Bathyplectes curculionis (Hymenoptera: Ichneumonidae).. Journal of Applied Entomology, 128, 316-320.
        • Evans, E.W, Richards, D.R, Kalaskar, A., (2004). Using foods for different purposes: female responses to prey in the predator Coccinellaseptempunctata L. (Coleoptera: Coccinellidae). Ecological Entomology, 29, 27-34.
        • Evans, E.W, (2003). Searching and reproductive behaviour of female aphidophagous ladybirds (Coleoptera: Coccinellidae): a review. European Journal of Entomology, 100, 1-10.
        • Rieder, J.P, Evans, E.W, Durham, S.L, (2001). Distribution of insect attacks in the biological control of weeds: infestation of Centaurea virgata flowerheads by a gall fly. Biological Control, 20, 254-260.
        • Kalaskar, A., Evans, E.W, (2001). Larval responses of aphidophagous lady beetles (Coleoptera: Coccinellidae) to weevil larvae versus aphids as prey. Annals of the Entomological Society of America, 94, 76-81.
        • Jacob, H.S, Evans, E.W, (2001). The influence of food deprivation on foraging decisions of the parasitoid Bathyplectes curculionis (Hymenoptera: Ichneumonidae). Annals of the Entomological Society of America, 94, 605-611.
        • Evans, E.W, (2000). Egg production in response to combined alternative foods by the predator Coccinella transversalis. Entomologia Experimentalis et Applicata, 94, 141-147.
        • Jacob, H.S, Evans, E.W, (2000). Influence of carbohydrate foods and mating on longevity of the parasitoid Bathyplectes curculionis (Hymenoptera: Ichneumonidae). Environmental Entomology, 29, 1088-1095.
        • Jacob, H.S, Evans, E.W, (2000). Influence of experience on the response of Bathyplectes curculionis (Hymenoptera: Ichneumonidae), a non-aphidophagous parasitoid, to aphid odor. Biological Control, 19, 237-244.
        • van der Werf, W., Evans, E.W, Powell, J., (2000). Measuring and modeling dispersal of Coccinella septempunctata in alfalfa fields. European Journal of Entomology, 97, 487-493.
        • van der Werf, W., Evans, E.W, Powell, J., (2000). Migration and dispersal of Coccinella septempunctata in Utah alfalfa. Proc., Experimental and Applied Entomology (Netherland Entomological Society, Am, 11, 135-144.
        • Evans, E.W, (2000). Morphology of invasion: body size patterns associated with establishment of Coccinella septemptunctata in western North America. European Journal of Entomology, 97, 469-474.
        • Evans, E.W, Stevenson, A.T, Richards, D.R, (1999). Essential versus alternative foods of insect predators: Benefits of a mixed diet. Oecologia, 121, 107-112.
        • Jacob, H.S, Evans, E.W, (1998). Effects of sugar spray and aphid honeydew on field populations of the parasitoid Bathyplectescurculionis (Hymenoptera: Ichneumonidae). Environ. Entomol., 27, 1563-1568.
        • Richards, D.R, Evans, E.W, (1998). Reproductive responses of aphidophagous lady beetles (Coleoptera: coccinellidae) to nonaphid diets: An example from alfalfa. Ann. Entomol. Soc. Am., 91, 632-650.
        • England, S., Evans, E.W, (1997). Effects of pea aphid (Homoptera: Aphididae) honeydew on longevity and fecundity of the alfalfa weevil (Coleoptera: Curculionidae) parasitoid Bathyplectes curculionis (Hymenoptera: Icheumonidae). Environmental Entomology, 26, 1437-1441.
        • Evans, E.W, , Richards, D.R, (1997). Managing the dispersal of ladybird beetles (Col.: Coccinellidae): Use of artificial honeydew to manipulate spatial distributions. Entomophaga, 42, 93-102.
        • Evans, E.W, England, S., (1996). Indirect interactions in biological control of insects: Pests and natural enemies in alfalfa. Ecological Applications, 6, 920-930.
        • Evans, E.W, (1995). Interactions among grasshoppers (Orthoptera: Acrididae) in intermountain grassland of western North America. Oikos, 73:73-78
        • Youssef, N.N, Evans, E.W, (1994). Exploitation of Canada Thistle by the weevil Rhinocyllus conicus (Coleoptera: Curculionidae) in northern Utah. Environmental Entomology, 23:1013-1019
        • Evans, E.W, Swallow, J.G, (1993). Numerical responses of natural enemies to artificial honeydew in Utah alfalfa. Environmental Entomology, 22:1392-1401
        • Evans, E.W, Bailey, K.W, (1993). Sampling grasshoppers (Orthoptera: Acrididae) in Utah grasslands: pan trapping versus sweep sampling. J. of the Kansas Entomological Soc., 66:214-222
        • Evans, E.W, (1992). Absence of interspecific competition among tallgrass prairie grasshoppers during a drought. Ecology, 73:1038-10
        • Evans, E.W, Youssef, N.N, (1992). Numerical responses of aphid predators to varying prey density among Utah alfalfa fields. J. of the Kansas Entomological Soc., 65:30-38
        • Evans, E.W, (1991). Experimental manipulation of herbivores in native tallgrass prairie: Response of aboveground arthropods. Amer. Midland Naturalist, 125:37-46
        • Evans, E.W, (1991). Intra versus interspecific interactions of ladybeetles (Coleoptera: Coccinellidate) attacking aphids. Oecologia, 87:401-408
        • Evans, E.W, (1990). Dynamics of an aggregation of blister beetles (Coleoptera:Meloidae) attacking a prairie legume. J. of the Kansas Entomological Society, 63:616-625
        • Evans, E.W, (1989). Interspecific interactions among phytophagous insects of tallgrass prairie: An experimental test. Ecology, 70:435-444
        • Evans, E.W, (1989). Timing of reproduction in a prairie legume: Seasonal impacts of insects consuming flowers and seeds. Oecologia, 78:220-230
        • Professional Journal

        • Evans, E.W, Karren, J.B, Israelsen, C.E, , (2007). Interactions over time between the cereal leaf beetle (Oulema melanopus, Coleoptera:Chrysomelidae)< and the larval parasitoid, Tetrastichus julis (Walker) (Hymenoptera:Eulophidae) in Utah.. Journal of Economic Entomology, 99, 1967-1973.
        • Public or Trade Journal

        • Evans, E.W, (1991). Alfalfa weevil has nemesis of its own. Utah Farmer-Stockman *
        • Evans, E.W, (1989). Lady beetles play key role as farmers' friends. Utah Farmer-Stockman, April *
        • Evans, E.W, (1989). Monitor alfalfa fields carefully before spraying for weevils. Utah Farmer-Stockman, April, Special Insert: 13 *
        • Evans, E.W, (1988). Explore alternatives to spraying alfalfa for insect control.. Utah Farmer-Stockman October Special Insert: 11 *

        * Has not been peer reviewed

        Publications - Literary Journal

          * Has not been peer reviewed

          Publications - MultiMedia

            * Has not been peer reviewed

            Publications - Technical Reports

              Other Reports

            • Evans, E.W, (2010). Dynamics and impact of Coccinella septempunctata as an invasive ladybird beetle in North America. IOBC/wprs Bulletin

            * Has not been peer reviewed

            Publications - Translations & Transcripts

              Publications - Other

                Other

              • Evans, E.W, (2008). Book review: Ladybugs of Alberta, by John Acorn.. American Entomologist *
              • Evans, E.W, Hodgson, E.W, (2008). Grasshoppers.. Utah Pests Fact Sheet (Ent-125-08) *
              • Hodgson, E.W, Evans, E.W, (2007). Cereal leaf beetle. Utah Pests Fact Sheet (Ent-84-07-PR) *
              • Evans, E.W, (2001). Expanding predator-prey theory for insects (Book review of A.F.G. Dixon, Insect predator-prey dynamics: ladybird beetles and biological control). Ecology *
              • Evans, E.W, (1993). Book review: Ecology and field biology (4th ed.), R.L. Smith. American Biology Teacher *
              • Evans, E.W, (1993). Biological weed control in Utah. The Weeder's Digest *
              • Evans, E.W, (1993). Biological control agents for Utah weeds: Ceutorrhynchus litura, a stem-mining weevil of Canada thistle. Utah Coop. Ext. Ser., Ext. Entomology Fact Sheet *
              • Evans, E.W, (1993). Biological control agents for Utah weeds: The knapweed seedhead gall flies. Utah Coop. Ext. Ser., Ext. Entomology Fact Sheet *
              • Evans, E.W, (1993). Biological control agents for Utah weeds: The musk thistle weevil Rhinocyllus conicus. Utah Coop. Ext. Ser., Ext. Entomology Fact Sheet *
              • Evans, E.W, (1993). Biological control of weeds through use of introduced insects. Utah Coop. Ext. Service, Extension Entomology Fact Sheet *
              • Evans, E.W, (1993). Pea aphid outbreaks associated with spraying for the alfalfa weevil in Utah. Utah Coop. Ext. Ser., Ext. Entomology Fact Sheet *
              • Evans, E.W, (1993). What are the indirect costs of pesticide use?. Utah Coop. Ext. Service, Ext. Entomology Fact Sheet *
              • Evans, E.W, (1992). Field evaluation of insecticides for control of alfalfa weevil larvae in Panguitch Valley, UT. Utah Coop. Ext. Service, Ext. Entomology Fact sheet *
              • Evans, E.W, (1991). Several insects promise to control weeds. Utah Science, Fall Issue *
              • Evans, E.W, (1990). Chemical and biological control of grasshoppers in Utah. Utah Cooperative Extension Service, Extension Entomology Fact Sheet *
              • Evans, E.W, (1990). Farmers should benefit as immigrant lady beetles join native species. Utah Science, Summer Issue *
              • Evans, E.W, (1990). Grasshoppers in Utah: General Biology. Utah Cooperative Extension Service, Extension Entomology Fact Sheet *
              • Evans, E.W, (1989). The Alfalfa Weevil in Utah. Utah Cooperative Extension Service, Extension Entomology Fact Sheet *
              • Evans, E.W, (1989). Book review: Ecology and our endangered life-suppprt systems, by E.P. Odum. American Biology Teacher *

              * Has not been peer reviewed

              Scheduled Teaching

              BIOL 4500, 6750 - Applied Entomology, Spring 2017

              BIOL 3220 - Field Ecology, Fall 2016

              BIOL 3220 - Field Ecology, Fall 2016

              BIOL 4500, 6750 - Applied Entomology, Spring 2016

              BIOL 3220 - Field Ecology, Fall 2015

              BIOL 3220 - Field Ecology, Fall 2015

              BIOL 4500,6750 - Applied Entomology, Spring 2015

              BIOL 3220 - Field Ecology, Fall 2014

              BIOL 3220 - Field Ecology, Fall 2014

              BIOL 4500 - Applied Entomology, Spring 2014

              BIOL 6750 - Topics in Biology, Spring 2014

              BIOL 3220 - Field Ecology, Fall 2013

              BIOL 3220 - Field Ecology, Fall 2013

              BIOL 4500 - Applied Entomology, Spring 2013

              BIOL 3220 - Field Ecology, Fall 2012

              BIOL 3220 - Field Ecology, Fall 2012

              BIOL 4500 - Applied Entomology, Spring 2012

              BIOL 6750 - Topics in Biology, Spring 2012

              BIOL 3220 - Field Ecology, Fall 2011

              BIOL 3220 - Field Ecology, Fall 2011

              BIOL 4500 - APPLIED ENTOMOLOGY, Spring 2011

              BIOL 3220 - Field Ecology, Fall 2010

              BIOL 3220 - Field Ecology, Fall 2010


              Graduate Students Mentored

              Byron Love, Biology, August 2010
              Samantha Willden, Biology 2017
              Joan Meiners, Biology 2016
              Virginia Jennings Bolshakova, Biology 2013
              Nina P. Louden, Biology 2010
              Chad Keyser, Biology 2010
              L. Nicole Davidson, Biology 2008
              Yukie Kajita, Biology 2008
              Katie Swoboda, Biology 2007
              Julie P. Rieder, Biology 2005
              Meenakshi Venkataraman, Biology 2004
              Trent Toler, Biology, August 1997 - May 2001
              Helen Jacob, Biology, August 1994 - May 2000
              Douglas Richards, Biology, August 1994 - May 1997
              Suzanne DiGiacomo, Biology, August 1992 - May 1996
              Staci England, Biology, August 1992 - May 1995
              Susan Geer, Biology, August 1988 - May 1993
              Wendy Halsey, Biology, August 1990 - May 1992